Problem is that you actually REDUCE the underbonnet flow if you reverse either of the vents - think o it like this - you have air coming in the front in LARGE quantities (through rad and main scoop). The only way out is down, this doesnt clear the upper rear area of hot air. The vents at the back and top of the bay extract the stagnant air to help keep electronics and IC cooler. If you reverse them they tend to push air in at the top-back which reduces air circulation and to a small extent reduces air in at the front. Teh amount of extra "cold" air in is negligible (In fact I'd be surprised if any went in at all), its the interruption of flow that will simply reduce the efficiency. You have to realise that Toyota have a wind tunnel and many engineers working out the best way to put them.
Also, if you extract the air over the car you reduce the under-car pressure, giving better stability at speed. Possibly to a near-zero change but still, its all adding up.